Symphoricarpos, commonly known as the snowberry, waxberry, or ghostberry 俗稱雪莓、楊梅或鬼莓 シンフォリカルポス、一般的にスノ

Symphoricarpos, commonly known as the snowberry, waxberry, or ghostberry, is a small genus of about 15 species of deciduous shrubs in the honeysuckle family, Caprifoliaceae. With the exception of the Chinese coralberry, S. sinensis, which is indigenous to western China, all species are native to North and Central America. The name of the genus is derived from the Ancient Greek words συμφορεῖν (sumphoreîn), meaning “to bear together“, and καρπός (karpós), meaning “fruit“. It refers to the closely packed clusters of berries the species produces. Snowberry is a resilient plant able to withstand a variety of conditions. Snowberry plants are most commonly found in forests, dry or moist openings, rocky hillsides or near riverbanks and streams.[5] They have been known to grow in a variety of soil types such as light sandy soil, medium loamy soil and heavier clay soil. Snowberry plants are also able to grow in a wide range of acidic and basic pHs and sunlight conditions. Symphoricarpos leaves are 1.5–5 cm (– in) long, rounded, entire or with one or two lobes at the base. The flowers are small, greenish-white to pink, in small clusters of 5–15 together in most species, solitary or in pairs in some (e.g. S. microphyllus). The fruit is conspicuous, 1–2 cm (0.5–1 in) in diameter, soft, varying from white (e.g. S. albus) to pink (S. microphyllus) to red (S. orbiculatus) and in one species (S. sinensis), blackish purple. When the white berries are broken open, the interior looks like fine, sparkling granular snow. The flesh is spongy and contains two 2–5 mm long, whitish stone seeds. The seeds, which contain endosperm and a small embryo, are egg-shaped and more or less flattened. They have a very tough, hard, impermeable covering, and so are very hard to germinate and may lie dormant for up to ten years. The white berries create a cracking sound when they are stepped into firm ground. Species accepted as of August 2015 Symphoricarpos acutus () Dieck – Sharpleaf snowberry: California, Nevada, Oregon ymphoricarpos albus (L.) – Common snowberry: Canada USA S. × chenaultii Rehder – Chenault coralberry S. × doorenbosii Krüssm. S. guadalupensis Correll – McKittrick’s snowberry: western Texas S. guatemalensis J.K. Williams: Guatemala S. hesperius – Trailing snowberry: California, Baja California, Oregon, Washington, British Columbia S. longiflorus – Desert snowberry: southwestern USA, northwestern Mexico S. microphyllus Kunth – Littleleaf snowberry: Mexico, Guatemala, New Mexico S. mollis Nutt. – Creeping snowberry: California S. occidentalis Hook. – Western snowberry, Wolfberry: USA Canada S. orbiculatus Moench – Coralberry: northeastern Mexico, eastern central USA S. oreophilus – Mountain snowberry: northern Mexico, western USA, western Canada S. palmeri – Palmer’s snowberry: Mexico, southwestern USA S. parishii Rydb.: California, Nevada, Baja California S. rotundifolius – Round-leaved snowberry: Mexico, southwestern USA S. sinensis – Chinese snowberry: China S. vaccinioides Rydb. – Roundleaf snowberry: California Snowberry is a hermaphroditic species meaning it contains both male and female reproductive organs. It has the ability to grow via seeds but typically reproduces by releasing shoots from a rhizome. This method of shoot dispersal allows snowberry to grow in dense populations of bushes and trees. Snowberry plants also tend to use a reproductive method called layering in which the plant’s vertical stems will wilt and droop until they touch surrounding soil. Upon making contact with soil, roots will begin to form. Snowberry plants are resilient and studies have proved they are able to tolerate dormant seasonal fires. These fires actually encourage the snowberry plant’s layering reproductive method, as the regeneration of new plants results in an increased number of stems and therefore more opportunities for layering to occur. Common snowberry (S. albus) is an important winter food source for quail, pheasant, and grouse, but is considered poisonous to humans. The berries contain the isoquinoline alkaloid chelidonine, as well as other alkaloids. Ingesting the berries causes mild symptoms of vomiting, dizziness, and slight sedation in children. Common snowberry is a popular ornamental shrub in gardens, grown for its decorative white fruit and wildlife gardening. It is also a useful landscaping plant due to its extreme versatility—tolerating sun, shade, heat, cold, drought, and inundation. Due to their low saponin content, snowberry was a common medicinal treatment used by several North American Indigenous tribes. Snowberry contains low concentrations of saponins, which are anti-carcinogenic and anti-inflammatory. Saponins have also been proven to help with immune function and decrease cholesterol. Saponins can be extremely toxic if consumed in excess, but was typically appli